Approximately 445 million years ago, with Gondwana positioned over the South Pole, accelerated CO₂ drawdown from volcanic rock weathering and organic carbon burial drove Earth from greenhouse to icehouse conditions. Continental ice sheets formed, sea levels dropped roughly 100 meters, and two extinction pulses eliminated approximately 85% of marine species. The first pulse resulted from rapid cooling; the second, shorter pulse from rapid warming caused anoxic water to flood continental shelves. Recent geochronological work establishes that the rate of temperature change, not its magnitude alone, controlled extinction intensity.