Archaeological sites in the lower Yangtze River region of eastern China, including Kuahuqiao and Hemudu in modern Zhejiang Province, reveal evidence of early rice cultivation by sedentary forager-cultivator communities. Analysis of rice spikelet bases shows a gradual transition from wild to domesticated Oryza sativa japonica over millennia. The Kuahuqiao community transformed local vegetation and managed cultivation environments, marking a phase in the domestication process that would eventually sustain billions of people.