Approximately 55.8 million years ago, a massive release of carbon raised global temperatures by 5 to 8 degrees Celsius over roughly 200,000 years. Ocean acidification dissolved carbonate shells of deep-sea benthic foraminifera, causing a severe deep-ocean extinction event. Simultaneously, mammalian species diversified and migrated across continents, with new orders appearing in North America and Europe. Atmospheric CO2 may have exceeded 2,000 parts per million during the peak warming interval.