Between 43,000 and 40,000 years ago, early modern humans crafted flutes from bird bones and mammoth ivory. Archaeologists discovered these instruments in Hohle Fels and Geißenklösterle caves in southern Germany's Swabian Jura. The flutes contained multiple finger holes positioned to produce a diatonic musical scale. Found alongside figurines and other artifacts, these instruments indicate that musical expression formed part of Upper Paleolithic culture and coincided with developments in symbolic behavior.